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Joe Paterno succeeded Engle as Penn State's head coach in 1966. Paterno, a former assistant coach under Engle, led the Nittany Lions to unprecedented success over his 46-year tenure, amassing a record of 409-136-3 and winning two national championships. He is widely regarded as one of the greatest college football coaches of all time, leaving an enduring legacy on Penn State football and college football as a whole.